Dubbed "The Werewolf" by the Gotham Press, he was the alter-ego of Anthony Romulus, a former successful Olympic athlete who sold himself into the service of Professor Milo for power and the means to maintain his prowess. Ingesting an experimental serum containing wolf DNA and hormones, he was granted enhanced strength and durability. However, the mutagen also caused Romulus to transform into a werewolf, losing all control and succumbing to his animalistic instincts.
After Milo betrays him and a confrontation with Batman that sees him plummeting into the river, he attempts to find a way to cure himself of the mutagen's effects.
Seeking the infamous geneticist Emile Dorian on his private island, he instead stumbled upon Dorian's creation Tygrus, pleading with him to help him find his "Father."
Despite his best efforts to keep his Werewolf persona in check, he eventually transformed in the middle of the streets of Gotham and went on a rampage. Tracking down Dorian in a private lab, he attempted to maul the scientist to death, only to be attacked by Man-Bat and Tygrus. The ensuing fight caused a lab explosion that seemingly killed them all. However, his body was never discovered by Batman during his investigation into the wreckage.
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